This article develops the thesis that the past quarter-century of electoral volatility in Germany reached a critical tipping point at the 2005 election. The two major parties of the Bonn Republic are now at their lowest combined share of the popular vote since the Federal Republic's founding in 1949. Electoral necessity and not, as in 1966, elite choice forced them into a grand coalition with little programmatic consensus. Their respective demographic cores-church-going Catholics for the CDU and unionized industrial workers for the SPD-have eroded as has the proportion of the electorate identifying with them. Institutional factors such as the electoral system have neither helped nor hindered these changes. The current grand coalition also faces a larger and more focused opposition than in 1966. The article concludes with some comparisons between the current German party system and its Italian counterpart of the late 1980s.